This fearsome behemoth is known as a Glutton.
It does not have a head structure at all, but instead simply a huge fissure in the shape of a flower bud. Strictly speaking, this could be classified as the beast's mouth, used for both attacking and devouring prey.
A Glutton's mouthparts look a lot like petals, constantly opening and closing. This is how Gluttons control the flow of tissue fluid in and out of their bodies. Once a Glutton is ready to spread Stardust, its mouthparts bloom, just like a real flower. Using a pressure-based mechanism, it then spews out a burst of concentrated Stardust into the air.
The Glutton also uses this method to attack and infect enemies.